BM, aero... I must admit this isn't as simple as I had thought.
What you are saying the forward translation of the plane is not tied to the speed/rotation of the wheels. Essentially, the wheels spin frictionlessly on the axle (assuming the axle is fixed to the aircraft). So, if the engines were off, and someone turned the treadmill on, the plane would just sit there because the wheels would counterspin to the treadmill. Do I understand you right?
Then if you turned the engines on, it would push the axle forwards, increasing the rpm of the wheels more, which would increase the speed of the treadmill. That makes sense.
Here's the BUT I can't square away in my head. Say the wheel has circumference of 2m and is spinning at 50 rpm. That's 100m/min. The treadmill is moving backwards at 100m/min. How has the wheel moved moved forwards wrt the treadmill? Because even if it is spinning frictionlessly on the axle, it is still on the axle. And if the wheel hasn't translated forward relative to the treadmill, then the axle can't have either.